The Gatsby Computational Neuroscience Unit is offering a 4-year PhD Programme in Theoretical Neuroscience and Machine Learning. The programme provides a unique opportunity for a critical mass of theoreticians to interact closely with one another, with the Sainsbury Wellcome Centre for Neural Circuits and Behaviour (SWC), with the Centre for Computational Statistics and Machine Learning (CSML), and with other research groups in related UCL departments. Students complete a 4-year PhD in either machine learning or theoretical neuroscience, with minor emphasis in the complementary field. Courses in the first year, taught with colleagues from the SWC and CSML, provide a comprehensive introduction to both fields and systems neuroscience. Students are encouraged to work and interact closely with researchers at the SWC and/or CSML to take advantage of this uniquely multidisciplinary research environment.